Publications and Products


cover of Diptera Diversity

Diptera Diversity: Status, Challenges and Tools

Edited by Thomas Pape, Daniel Bickel, and Rudolf Meier

This new 460 page book was edited and features chapters by many of the FLYTREE principal investigators and collaborators. Set for release at the end of March 2009, this long-awaited book grew out of the 2002 International Congress of Dipterology in Brisbane, Australia. It features sections on regional diversity of Diptera fauna; case studies, ecological approaches & estimation; and bioinformatics and dipteran diversity (pdf of table of contents).

Evolutionary Biology of Flies

The Evolutionary Biology of Flies

David K. Yeates & Brian M. Wiegmann, eds.

“Flies have been at the heart of profound biological discovery. They are structurally and ecologically more diverse than all other orders of insects; they are paradigm organisms in genetics and physiology; they are the most effective vectors of diseases; and they have been the focus of revolutionary thinkers like Willi Hennig. This book integrates for the first time the remarkable biology of flies and brings the study of these insects to a new level. “

— David Grimaldi, American Museum of Natural History, author ofEvolution of Insects

“By so smoothly covering every aspect of the evolutionary biology of flies, this book certainly lives up to its title.”

— Donald A. Windsor, Science Books & Films

Preview in GoogleBooks

Finding the Fly Tree of Life – Poster: Isaac Winkler, North Carolina State University, Sept. 2011 (ver. 4)


Peer Reviewed Publications*

Jump to:
Comparative Genomics
Fossil Diptera
Fly Biology, Ecology, & Biodiversity
Dipteran Taxonomy
Fly Phylogeny & Evolution
Fly Morphology & Comparative Anatomy
Tools for Documenting Diptera Diversity

Comparative Genomics

Cameron, S. L., C. L. Lambkin, S. C. Barker, and M. F. Whiting. 2007. A mitochondrial genome phylogeny of Diptera: whole genome sequence data accurately resolve relationships over broad timescales with high precision.Systematic Entomology, 32: 40-59. (pdf(cited by journal as one of the most downloaded articles of 2008!)

Cameron, S. L., A. T. Beckenbach, M. A. Dowton and M. F. Whiting. 2006. Insect interordinal phylogenetics and mitochondrial genomics. Arthropod Systematics and Phylogeny. 64: 27-34. (pdf)

Hare, E. E., Peterson, B. K., Iyer V. N., Meier, R., Eisen, M. B. 2008. Sepsid even-skipped enhancers are functionally conserved in Drosophila despite lack of sequence conservation. PLoS Genetics 4: e1000106. (abstract) (pdf)

Meier, R. 2008. DNA Sequences in Taxonomy: Opportunities and Challenges. Pp. 95-128. in Wheeler QD (ed.),The New Taxonomy Systematics Association Special Volume. CRC Press. (preview)

Meier, R. and G. Zhang. 2009. DNA barcoding and DNA taxonomy in Diptera: An assessment based on 4,261 COI sequences for 1,001 species. Pages 349-380 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Meier, R., G. Zhang, F. Ali. 2008. The use of mean instead of smallest interspecific distances exaggerates the size of the “barcoding gap” and leads to misidentification. Systematic Biology, 57: 809-813. (pdf)



Fossil Diptera

Amorim, D.D.S. and D. A. Grimaldi. 2006. Valeseguyidae, a new family of Diptera in the Scatopsoidea, with a new genus in Cretaceous amber from Myanmar. Systematic Entomology: 31: 508-516. (pdf 9.4 MB) (abstract)

Blagoderov, V.,  D. A. Grimaldi, and N. C. Fraser. 2007. How time flies for flies: Diverse Diptera from the Triassic of Virginia and early radiation of the order. American Museum Novitates (3572): 1-39. (pdf 10 MB)

Wiegmann, B. M., D. K Yeates, J. L. Thorne, and H. Kishino. 2003. Time flies, a new molecular time-scale for brachyceran fly evolution without a clock. Systematic Biology, 52: 745-756. (pdf, 655 KB)

Yeates, D. K., R. Meier, and B. M. Wiegmann. 2003. Phylogeny of true flies (Diptera): A 250 million year old success story in terrestrial diversification. Pages 170-172 in Klass, K.-D. ed. Proceedings of the 1st Dresden Meeting on Insect Phylogeny: “Phylogenetic Relationships within the Insect Orders” (Dresden, September 19-21, 2003).Entomologische Abhandlungen, 61: 119-172. (pdf)



Fly Biology, Ecology, & Biodiversity

Amorim, D.D.S. 2009. Neotropical Diptera diversity: Richness, patterns, and persectives. Pages 71-97 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Bänziger, H. & Pape, T. 2004. Flowers, faeces and cadavers: natural feeding and laying habits of flesh flies in Thailand (Diptera: Sarcophagidae, Sarcophaga spp.). – Journal of Natural History 38: 1677-1694. (article)

Bickel, D. 2009. Biogeography of Diptera in the Southwest Pacific. Pages 257-278 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Cerretti, P. & Pape, T. 2004. Metopodia pilicornis (Pandellé, 1895) kleptoparasitic on Sphex funerariusGussakovskij, 1934 (Diptera: Sarcophagidae, Miltogramminae; Hymenoptera: Sphecidae). Studia dipterologica 10: 557-558. (table of contents)

Chaverri, L. G., and A. Borkent. 2007. The Meniscus midges of Costa Rica (Diptera: Dixidae). Zootaxa, 1575: 1-34. (pdf) (abstract)

Courtney, G.W.Pape, T., Skevington, J.H. and Sinclair, B.J. 2009. Biodiversity of Diptera. Pp. 183-221 in: Foottit, R.G. & Adler, P.H. (eds), Insect Biodiversity: Science and Society. Blackwell Publishing, Oxford.

Dupont, S. & Pape, T. Termitophile and termite-associated scuttle flies (Diptera: Phoridae). Terrestrial Arthropod Reviews [In press.]

Evenhuis, N. L. 2009. Hawaii’s Diptera biodiversity. Pages 47-70 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Ferguson, D. J. 2007. Field observations of Perissomma mcalpinei Colless (Diptera: Perissommatidae). Australian Entomologist, 34: 93-96.

Grootaert, P. 2009. Oriental Diptera, a challenge in diversity and taxonomy. Pages 197-226 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Hagman, M., Pape, T. & Schulte, R. 2005. Flesh fly myiasis (Diptera: Sarcophagidae) in Peruvian poison frogs genus Epipedobates (Anura, Dendrobatidae). – Phyllomedusa 4: 69-73. (pdf)

Kirk-Spriggs, A. H. and B. R. Stuckenberg. 2009. Afrotropical Diptera — Rich savannas, poor rainforests. Pages 155-196 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Méndez, J., Mello-Patiu, C. de & Pape, T. 2008. New flesh flies (Diptera: Sarcophagidae) from coastal mangroves of Panama. – Journal of Natural History 42: 249-257. (article)

Pape, T. 2009. Palaearctic Diptera — from Tundra to Desert. Pages 121-154 inPape, T., Bickel, D. & Meier, R.(eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Pape, T., Bickel, D. & Meier, R. (eds) 2009. Diptera Diversity: Status, Challenges and Tools. Brill, Leiden, 459 pp.

Pape, T., Thompson, F.C. & Szpila, K. 2008. What’s in a frog stomach? Solving a 150 year old mystery (Diptera: Calliphoridae). – Systematic Entomology 33: 548-551. (pdf)

Pollet, M. 2009. Diptera as ecological indicators of habitat and habitat change. Pages 302-322 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Sinclair, B. J. 2009. Dipteran biodiversity of the Galápagos. Pages 98-120 in: Pape, T., Bickel, D. & Meier, R.(eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Ssymank, A., Kearns, C.A., Pape, T. & Thompson, F.C. 2008. Pollinating flies (Diptera): a major contribution to plant diversity and agricultural production. – Biodiversity 9: 86-89. (table of contents)

Stuke, J.-H. and J. H. Skevington. 2007. The Conopidae of Costa Rica (Diptera) (Part 1: Conopinae – Conopini &Tropidomyiini). Zootaxa, 1528: 1-40. (pdf) (abstract)

Sze, W.T., Pape, T. & O’Toole, D.K. 2008. The first blow fly parasitoid takes a head start in its termite host (Diptera: Calliphoridae, Bengaliinae; Isoptera: Macrotermitidae). Systematics and Biodiversity 6: 25-30. (abstract may access full article from this link with subscription)

Thompson, F. C. 2009. Nearctic Diptera: Twenty years later. Pages 3-46 inPape, T., Bickel, D. & Meier, R.(eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Whitmore, D., Richet, R. & Pape, T. 2008. Sarcophagidae. In: Ziegler, J. (ed.), Diptera Stelviana. A dipterological perspective on a changing alpine landscape. – Studia Dipterologica Supplement 16: 229-237.

Yeates, D. K., D. Bickel, D. K. McAlpine, and D. H. Colless. 2009. Diversity, relationships and biogeography of Australian flies. Pages 227-256 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.



Dipteran Taxonomy

Amorim, D.D.S. and M.I.P.A. Balbi. 2006. A review of Anapausis Enderlein (Diptera: Scatopsidae) in the Neotropical Region, with four new species and comments on the phylogeny of the genus. Zootaxa, 1300: 1-29. (pdf) (abstract)

Ang, Y., G. S. Lim, R. Meier. 2008. Morphology and DNA sequences confirm the first Neotropical record for the Holarctic sepsid species Themira leachi (Meigen) (Diptera: Sepsidae). Zootaxa, 1933: 63-65. (abstract & excerpt) (pdf *)

Ang, Y., N. Puniamoorthy, and R. Meier. 2008. Secondarily reduced foreleg armature in Perochaeta dikowi sp. n. (Diptera: Cyclorrhapha: Sepsidae) due to a novel mounting technique. Systematic Entomology, 33: 552-559. (abstract) (pdf)

Barraclough, D. A. 2006. An overview of the South African tangle-veined flies (Diptera: Nemestrinidae), with an annotated key to the genera and a checklist of species. Zootaxa, 1277: 39-63. (pdf) (abstract)

Barraclough, D. A. 2006. Remarkable sexual dimorphism in the Afrotropical species Nycterimyia capensis Bezzi, 1924 (Diptera: Nemestrinidae), and its taxonomic implications. Zootaxa, 1273: 1-7. (pdf) (abstract)

Bickel, D. 2009. Why Hilara is not amusing: the problem of open-ended taxa and the limits of taxonomic knowledge. Pages 279-301 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Blackith, R.M., Richet, R., and Pape, T. 2004. Revision of Sarcophaga dissimilis Meigen, 1826 and Sarcophaga chaetoneura Brauer & Bergenstamm, 1889 (Diptera: Sarcophagidae). – Zootaxa 560: 1-13. (pdf)

Bortolanza, M., C.J.B. De Carvalho, and A.P.C. Lara. 2006. A new species of Coenosopsia Malloch (Diptera, Anthomyiidae) from southern Brazil and a reappraisal of the cladistic relationships of the genus. Zootaxa, 1242: 37-52. (pdf) (abstract)

Brake, I. and W. N. Mathis. 2006. Revision of the genus Australimyza Harrison (Diptera: Australimyzidae).Systematic Entomology: 32: 252-275. (pdf) (abstract)

Cerretti, P. & Pape, T. 2007. Two new species of European Stevenia Robineau-Desvoidy (Diptera: Rhinophoridae) and a key to the Palaearctic species. – Zootaxa 1624: 31-41. (pdf)

De CarvalhoC.J.B. and A.C. Pont. 2006. Taxonomy, cladistics and biogeography of the South American genusBrachygasterina Macquart (Diptera: Muscidae). Zootaxa, 1151: 1-26. (pdf) (abstract)

Evenhuis, N.L., Pape, T. & Pont, A.C. 2008. The problems of subsequent typification in genus-group names and use of the Zoological Record: a study of selected post-1930 Diptera genus-group names without type species designations. – Zootaxa 1912: 1-44. (abstract) (pdf *)

Holston, K. C. and M. E. Irwin. 2005. Revision of the Nearctic Thereva (Diptera, Asiloidea: Therevidae). Studia dipterologica Supplement 13, 219 pp. (summary)

Lonsdale, O. and S. A. Marshall. 2006. Revision of the New World species of Craspedochaeta (Diptera: Clusiidae).Zootaxa, 1291: 1-101. (pdf) (abstract)

Meier, R. 2007. Themira biloba Andersson 1975 (Sepsidae: Diptera), a species from Manhattan’s Central Park that is new to the Nearctic Region. Journal of the New York Entomological Society, 114: 176-177. (pdf)

Nihei, S. S. and C.J.B. De Carvalho. 2006. Systematics and biogeography of Polietina Schnabl & Dziedzicki (Diptera, Muscidae): Neotropical area relationships and Amazonia as a composite area. Systematic Entomology: 32: 477-501. (pdf) (abstract)

Otranto, D., Colwell, D.D. and Pape, T. 2005. Hypoderma sinense Pleske, 1926: solving a century-old enigma.Medical and Veterinary Entomology 19: 315-321. (abstract)

Pape, T. 2004. The Sarcophagidae (Insecta: Diptera) described by Louis Pandellé. – Zootaxa 722: 1-64. (abstract & excerpt) (pdf *)

Pape, T. 2006. A new species of Hoplacephala Macquart (Diptera: Sarcophagidae) from Namibia, with a discussion of generic monophyly. Zootaxa, 1183: 57-68. (pdf *) (abstract)

Pape, T. 2008. Nominal species of Australasian/Oceanian Sarcophagidae proposed by J. Macquart (Insecta: Diptera). – Zootaxa 1702: 41-51. (abstract & excerpt) (pdf *)

Pape, T. and C. A. De Mello-Patiu. 2006. Revision of Engelimyia Lopes, 1975 (Diptera: Sarcophagidae). Zootaxa,1256: 21-47. (pdf) (abstract)

Pape, T. and Kurahashi, H. 2004. Three new species of Sarcophaga (Insecta: Diptera: Sarcophagidae) from Timor, with a lectotype designation for Myophora peroniiSpecies Diversity 9: 165-185. (abstract)

Perez-Gelabert, D. E. and F. C. Thompson. 2006. A new genus and species of Richardiidae (Diptera) from Hispaniola. Zootaxa, 1259: 25-31. (pdf) (abstract)

Petersen, F. T., J. Damgaard, and R. Meier. 2007. DNA Taxonomy: How many DNA sequences are needed for solving a taxonomic problem? The case of two parapatric species of louse flies (Diptera: Hippoboscidae:Ornithomya (Latreille, 1802). Arthropod Systematics & Phylogeny, 65: 111-117. (pdf)

Santos, D.M.D. and D. S. Amorim. 2007. Chrysopilus (Diptera: Rhagionidae) from Brazil: redescription ofChrysopilus fascipennis Bromley and description of eleven new species. Zootaxa, 1510: 1-33. (pdf) (abstract)

Szpila, K. and Pape, T. 2007. Rediscovery, redescription and reclassification of Beludzhia phylloteliptera (Diptera: Sarcophagidae, Miltogramminae). European Journal of Entomology 104: 119-137. (abstract) (pdf)

Whitmore, D., Richet, R., Pape, T. and Blackith, R.M. 2009. Redescription of Sarcophaga (DiscachaetabezzianaBöttcher and Sarcophaga (Heteronychiainfixa Böttcher, and description of a new Heteronychia Brauer & Bergenstamm from southern France (Diptera: Sarcophagidae). Zootaxa 1993: 27-40. (pdf *)

Winterton, S. L. 2007. New species of Nanexila Winterton & Irwin and Taenogera Krober from Australia (Diptera: Therevidae). Zootaxa, 1413: 55-64. (pdf) (abstract)

Winterton, S. L. 2007. New species of Acraspisoides Hill & Winterton and Bonjeania Irwin & Lyneborg (Diptera: Therevidae: Agapophytinae), with the description of a new genus. Zootaxa, 1438: 1-25. (pdf) (abstract)

Winterton, S. L. 2007. New species of Laxotela Winterton & Irwin from Australia (Diptera: Therevidae: Agapophytinae). Zootaxa, 1407: 57-64. (pdf) (abstract)

Winterton, S. L. and P. H. Kerr. 2006. A new species of Alloxytropus Bezzi (Diptera: Scenopinidae: Proratinae) from Israel. Zootaxa, 1155:41-50. (pdf) (abstract)

Winterton, S. L. 2007. Revision of the Australian stiletto fly genus Acatopygia Krober (Diptera: Therevidae: Agapophytinae). Zootaxa, 1405: 51-62. (pdf) (abstract)

Winterton, S. L. 2006. New species of Eupsilocephala Krober from Australia (Diptera: Therevidae). Zootaxa, 1372: 17-25. (pdf) (abstract)

Yeates, D. K., M. E. Irwin, and B. M. Wiegmann. 2003. Ocoidae, a new family of asiloid flies (Diptera: Brachycera: Asiloidea), based on Ocoa chilensis gen. and sp.n. from Chile, South America. Systematic Entomology, 28: 417-431. (pdf) (abstract)

Yeates, D. K., M. E. Irwin, and B. M. Wiegmann. 2006. Evocoidae (Diptera: Asiloidea), a new family name for Ocoidae, based on Evocoa, a replacement name for the Chilean genus Ocoa Yeates, Irwin, and Wiegmann 2003.Systematic Entomology, 31: 373. (pdf) (Summary)



Fly Phylogeny and Evolution

Amorim, D. S. and  D. Yeates. 2006. Pesky Gnats: Ridding Dipteran Classification of the Nematocera. Studia Dipterologica, v. 13, p. 3-9. (abstract)

Amorim, D. S. and E. Rindal. 2007.  Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535: 1-92. (pdf) (abstract)

Beckenbach, A. T., and A. Borkent. 2003. Molecular analysis of the biting midges (Diptera: Ceratopogonidae), based on mitochondrial cytochrome oxidase subunit 2. Molecular Phylogenetics and Evolution, 27: 21-35. (pdf) (abstract)

Bertone M. A., G. W. Courtney and B. M. Wiegmann. 2008. Phylogenetics and a timescale for diversification of the earliest true flies (Insecta: Diptera) based on multiple nuclear genes. Systematic Entomology. 33: 668-687.

Bertone, M. A., and B. M. Wiegmann. 2009. Diptera. In: S. Kumar and S. B. Hedges, eds. The Timetree of Life. Oxford University Press. in press.

Bickford, D., Lohman, D., Sodhi, N. S., Ng, P. K. L., R. Meier, K. Winker, K. Ingram, I. Das. 2006. Cryptic species: a new window on diversity and conservation. Trends in Ecology and Evolution, 22: 148-155. (pdf) (Summary)

Doczkal, D. & Pape, T. Lyneborgimyia magnifica gen. et spec. nov. (Diptera: Syrphidae) from Tanzania, with a phylogenetic analysis of the Eumerini using new morphological characters. – Systematic Entomology [In press.]

Gibson, J. F. and G. W. Courtney. 2006. Revision of the net-winged midge genus Horaia Tonnoir and its phylogenetic relationship to other genera within the tribe Apistomyiini (Diptera: Blephariceridae). Systematic Entomology, 32: 276-304. (pdf) (abstract)

Giroux, M., Pape, T. & Wheeler, T.A. Towards a phylogeny of the flesh flies (Diptera: Sarcophagidae): morphology and phylogenetic implications of the acrophallus in the subfamily Sarcophaginae. – Zoological Journal of the Linnean Society [In press.]

Holston, K. C., B. M. Wiegmann, and M. E. Irwin. 2007. Monophyly and phylogenetic relationships of Therevaand therevine genus groups (Insecta: Diptera: Therevidae) based on EF-1a, 28S rDNA, and mt16S rDNA sequences. Invertebrate Systematics, 21: 279-296. (abstract)

Ingram, K. K., T. Laamanen, N. Puniamoorthy, and R. Meier. 2008. Lack of morphological coevolution between male forelegs and female wings in Themira (Sepsidae: Diptera: Insecta). Biological Journal of the Linnean Society, 93: 227-238. (abstract) (pdf)

Kutty, S. N., M. V. Bernasconi, F. Sifner, and R. Meier. 2006. Sensitivity analysis, molecular systematics, and natural history evolution of Scathophagidae (Diptera: Cyclorrhapha: Calyptratae). Cladistics, 23: 64-83. (pdf) (abstract)

Kutty, S.N., Pape, T., Pont, A.C., Wiegmann, B.M. and Meier, R. 2008. The Muscoidea (Diptera: Calyptratae) are paraphyletic: Evidence from four mitochondrial and four nuclear genes. – Molecular Phylogeny and Evolution49: 639-652. (article)

Laamanen, T. R., R. Meier, M. A. Miller, A. Hille, and B. M. Wiegmann. 2005. Phylogenetic analysis of Themira(Sepsidae: Diptera): sensitivity analysis, alignment, and indel treatment in a multigene study. Cladistics, 21:258-271. (pdf) (abstract)

Lim, G. S., W. S. Hwang, S. N. Kutty, R. Meier, and P. Grootaert 2009. Mitochondrial and nuclear markers support the monophyly of Dolichopodidae and suggest a rapid origin of the subfamilies (Diptera: Empidoidea). Systematic Entomology [in press].

Marinoni, L. and W. N. Mathis. 2006. A cladistic analysis of the Neotropical genus Sepedonea Steyskal (Diptera: Sciomyzidae). Zootaxa, 1236: 37-52. (pdf) (abstract)

Meier, R. 2005. The role of Dipterology in phylogenetic systematics: Insights of Willi Hennig. Pp. 45-62 in Yeates, D. K. and B. M. Wiegmann (eds), The Evolutionary Biology of Flies. Columbia University Press, New York. (excerpt)

Meier, R., S. Kwong, G. Vaidya, and P.K.L. Ng. 2006. DNA Barcoding and taxonomy in Diptera: a tale of high intraspecific variability and low identification success. Systematic Biology, 55: 715-728. (abstract) (pdf)

Moulton, J. K. 2003. Can the current molecular arsenal adequately track rapid divergence events within Simuliidae (Diptera)? Molecular Phylogenetics and Evolution, 27: 45-57. (pdf) (abstract)

Moulton, J. K. and B. M. Wiegmann. 2003. Evolution and phylogenetic utility of CAD (rudimentary) among Mesozoic-aged eremoneuran Diptera (Insecta). Molecular Phylogenetics and Evolution, 31: 363-378. (pdf)

Moulton, J. K. and B. M. Wiegmann. 2007. The phylogenetic relationships of flies in the superfamily Empidoidea (Insecta: Diptera). Molecular Phylogenetics and Evolution, 43: 701-713. (pdf)

Nihei, S. S. and C.J.B. De Carvalho. 2004. Taxonomy, cladistics and biogeography of Coenosopsia Malloch (Diptera, Anthomyiidae) and its significance to the evolution of anthomyiids in the Neotropics. Systematic Entomology: 29: 260-275. (pdf) (abstract)

Pape, T. 2006. Phylogeny and evolution of the bot flies. Pages 20-50 in: Colwell, D., Scholl, P. & Hall, M. (eds), The Oestrid Flies: biology, host-parasite relationships, impact and management. CABI Publishers; Wallingford and Cambridge, MA.

Papoucheva, P., V. Proviz, C. Lambkin, B. Goddeeris and A. Blinov. 2003. Phylogeny of the endemic BaikalianSergentia (Chironomidae, Diptera). Molecular Phylogenetics and Evolution, 29: 120-125. (pdf) (abstract)

Petersen, F. T., R. Meier, S. Kutty, and B. M. Wiegmann. 2007. The phylogeny and evolution of host choice in the Hippoboscoidea (Diptera) as reconstructed using four molecular markers. Molecular Phylogenetics and Evolution, 45: 111-122. (pdf)

Puniamoorthy, N., K. Feng Yi Su, R. Meier. 2008. Bending for love: losses and gains of sexual dimorphisms are strictly correlated. with changes in the mounting position of sepsid flies (Sepsidae: Diptera). BMC Evolutionary Biology, 8:155. (abstract) (pdf)

Savage, J., T. A. Wheeler, and B. M. Wiegmann. 2004. Phylogenetic analysis of the genus Thricops Rondani (Diptera: Muscidae) based on molecular and morphological characters. Systematic Entomology, 29: 395-414. (pdf2.1 MB)

Scheffer, S. J. 2005. Invasive Diptera: Using molecular markers to investigate cryptic species and the global spread of introduced flies. Pp. 371-387 in Yeates, D. K. and B. M. Wiegmann (eds), The Evolutionary Biology of Flies. Columbia University Press, New York. (excerpt)

Scheffer, S. J., M. L. Lewis, R. C. Joshi. 2006. DNA barcoding applied to invasive leafminers (Diptera: Agromyzidae) in the Philippines. Annals of the Entomological Society of America. 99: 204-210.

Scheffer, S. J., I. S. Winkler, and B. M. Wiegmann. 2007. Phylogenetic relationships within the leaf-mining flies (Diptera: Agromyzidae) inferred from sequence data from multiple genes. Molecular Phylogenetics and Evolution,42: 756-775. (pdf)

Schnell e Schuehli, G., C.J.B. Carvalho, and B. M. Wiegmann. 2007. Molecular phylogenetics of the Muscidae: New ideas in a congruence context. Invertebrate Systematics, 21: 263-278. (abstract).

Sinclair, B.J., A. Borkent and D. M. Wood. 2007.  The male genital tract and aedeagal components of the Diptera with a discussion of their phylogenetic significance. Zoological Journal of the Linnaean Society, 150: 711-742. (abstract) (pdf *)

Skevington, J. H., C. Kehlmaier, and G. Stahls. 2007 DNA Barcoding: Mixed results for big-headed flies (Diptera: Pipunculidae). Zootaxa, 1423: 51-57. (pdf)

Stevens, J.R., Wallman, J.F., Otranto, D., Wall, R. & Pape, T. 2006. The evolution of myiasis in humans and other animals in the Old and New Worlds (part II): biological and life-history studies. Trends in Parasitology 22: 181-188. (article)

Wiegmann, B. M., J. Kim and M. D. Trautwein. 2009. Holometabola. In: S. Kumar and S. B. Hedges, eds. The Timetree of Life. Oxford University Press. in press.

Wiegmann, B. M., M D. Trautwein, J. Kim, M. Bertone, S. L. Winterton, B. K. Cassel, and D. K. Yeates. 2009. Nuclear genes resolve phylogeny of the holometabolous insect orders. BMC Evolutionary Biology, in press.

Winkler, I. S., S. J. Scheffer, and C. Mitter. 2009. Molecular phylogeny and systematics of leaf-mining flies (Diptera: Agromyzidae): delimitation of Phytomyza Fallén sensu lato and included species groups, with new insights on morphological and host-use evolution. Systematic Entomology 34(2): 260-292. (abstract) (pdf) (supporting information)

Winterton, S. L., B. M. Wiegmann, and E. I. Schlinger. 2007. Phylogeny and Bayesian divergence time estimations of small-headed flies (Diptera: Acroceridae) using multiple molecular markers. Molecular Phylogenetics and Evolution, 43: 808-832. (pdf)

Yeates, D. K. and B. M. Wiegmann (eds). 2005. The Evolutionary Biology of Flies. Columbia University Press, New York, 440 pp. (excerpts)

Yeates, D. K. and B. M. Wiegmann. 2005. Phylogeny and evolution of Diptera: Recent insights and new perspectives. Pp. 14-44 in Yeates, D. K. and B. M. Wiegmann (eds), The Evolutionary Biology of Flies. Columbia University Press, New York. (excerpt)

Yeates, D. K., Wiegmann, B. M., Courtney, G. W., Meier, R., Lambkin, C. and Pape, T. 2007. Phylogeny and systematics of Diptera: Two decades of progress and prospects. – Zootaxa 1668: 565-590. (pdf)



Fly Morphology & Comparative Anatomy

Dupont, S. and Pape, T. 2007. Fore tarsus attachment device of the male Aenigmatias lubbockii (Verrall, 1877).Journal of Insect Science 7(54): 1-8. (article)

Szpila, K. and Pape, T. 2005. Comparative morphology of the first instar of three species of Metopia Meigen (Diptera: Sarcophagidae, Miltogramminae). Acta Zoologica 86: 119-134. (abstract)

Szpila, K. and Pape, T. 2005. The first instar larva of Apodacra pulchra (Diptera: Sarcophagidae, Miltogramminae).Insect Systematics and Evolution 36: 293-300.

Szpila, K. and Pape, T. 2008. Morphological diversity of first instar larvae in Miltogramma subgenus Pediasiomyia(Diptera: Sarcophagidae, Miltogramminae). Zoologischer Anzeiger 247: 259-273. (article)

Szpila, K. and Pape, T. 2008. Morphology of the first instar larva of the housefly Musca domestica Linnaeus, 1758 (Diptera: Muscidae). Journal of Medical Entomology 45: 594-599. (article)

Szpila, K., Pape, T. and Rusinek, A. 2008. Morphology of the first instar larva of Calliphora vicinaPhormia reginaand Lucilia illustris (Diptera, Calliphoridae). Medical and Veterinary Entomology 22: 16-25. (abstract)



Tools for Documenting Diptera Diversity

Dikow, T., R. Meier, G. G. Vaidya, and J.G.H. Londt. 2009. Biodiversity research based on taxonomic revisions — a tale of unrealized opportunities. Pages 323-380 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden.

Kampmeier, G. E. and M. E. Irwin. 2009. Meeting the interrelated challenges of tracking specimen, nomenclature, and literature data in Mandala. Pages 407-437 inPape, T., Bickel, D. & Meier, R. (eds), Diptera Diversity: Status, Challenges and Tools. Brill, Leiden. (pdf available by permission from publisher)

Winterton, S. L. 2009. Diptera biodiversity informatics. Pages 381-406 inPape, T., Bickel, D. & Meier, R. (eds),Diptera Diversity: Status, Challenges and Tools. Leiden: Brill Academic Publishers, pp. 381-406.


Project Progress Reports

Year 1 Report – Year 2 Report – Year 3 Report – Year 4 Report – Year 5 Report


Yeates, D.K., J. Hamilton, and A. Hastings. Glossary of fly external morphology

Yeates, D., R. Meier, and B. Wiegmann.  Phylogenetic Supertree of 151 families of Diptera

Hamilton, J.R., D.K. Yeates, A. Hastings, D.H. Colless, D.K. McAlpine, D. Bickel, P.S. Cranston, M.A. Schneider, G. Daniels, S. Marshall. 2006. On the Fly – The Interactive Atlas and Key to Australia Fly Families. Centre for Biological Information Technology, CSIRO. [to obtain] [Winner of Whitley Book Award]

Workshops & Meetings

Tangalooma Diptera Workshops, 23-25 August 2004.

Copenhagen meeting


2006 International Congress of Dipterology, Fukuoka, Japan

2007 collective buzz at Coweeta, North Carolina 17-18 November 2007

2008 BioSynC meeting at the Field Museum, Chicago, IL 6-9 October


Entomological Society of America

Collecting Trips

California – Epistomia